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@optimuspig
I bought 1,500 sparks using 600 gems just for the heck of it. You can only purchase it once (although you can also buy 500 sparks for 210 gem/420 tokens and 750 sparks for 325 gems/750 tokens).
In the barracks it looks like the minimum spend is 500 sparks to get a 5% upgrade (whatever that is) on one character.